Dr. Jayme L. Waldron, Professor

Department of Biological Sciences
Marshall University
One John Marshall Drive
Huntington, WV 25755
[email protected]
Ph.D., Forest Resources, Clemson University, 2005
M.S., Biological Sciences, Marshall University, 2000
B.S., Wildlife & Fisheries Resources, West Virginia University, 1998
Courses Taught:
Herpetology (BSC 405/506)
Ornithology (BSC 408/508)
Conservation Biology (BSC 460/560)
Quantitative Ecology (BSC 644)
Graduate Seminar (BSC 662)
Marshall University
One John Marshall Drive
Huntington, WV 25755
[email protected]
Ph.D., Forest Resources, Clemson University, 2005
M.S., Biological Sciences, Marshall University, 2000
B.S., Wildlife & Fisheries Resources, West Virginia University, 1998
Courses Taught:
Herpetology (BSC 405/506)
Ornithology (BSC 408/508)
Conservation Biology (BSC 460/560)
Quantitative Ecology (BSC 644)
Graduate Seminar (BSC 662)
Research Interests
I am interested in questions about the application of life history data to wildlife conservation. I am particularly interested in using habitat and population models in the development of management and restoration strategies. Herpetofauna are model organisms for conservation studies because their life history attributes, e.g., reproductive strategies, age at reproductive maturity, longevity, and growth, tend to vary greatly within taxa, providing unique opportunities for comparative research that examines intra- and interspecific differences in species vulnerability to extinction.
I am interested in questions about the application of life history data to wildlife conservation. I am particularly interested in using habitat and population models in the development of management and restoration strategies. Herpetofauna are model organisms for conservation studies because their life history attributes, e.g., reproductive strategies, age at reproductive maturity, longevity, and growth, tend to vary greatly within taxa, providing unique opportunities for comparative research that examines intra- and interspecific differences in species vulnerability to extinction.
Publications (* denotes student author)
Edwards, E.*, B. Mosher, T.K. Pauley, S.M. Welch, and J.L. Waldron. In Review. The effect of liming on spring salamander abundance.
Journal of Wildlife Management.
Kelley, A.*, S. M. Welch, J. Holloway, J.W. Dillman, A. Atkinson, and J. L. Waldron. 2022. Effectiveness of long-distance translocation of
Eastern Diamondback Rattlesnakes. Wildlife Society Bulletin e1291. Https://doi.org/10.1002/wsb.1291.
Jungen, M.*, Z. Ross*, J. Cooley*, M. Martin, J. Holloway, S. M. Welch, and J. L. Waldron. 2019. Monitoring eastern diamondback
rattlesnakes using a novel external radio-transmitter attachment method. Copeia 107:411-416.
Amspacher, K.*, B. Bauer, J. Waldron, E. Wiggers, and S. Welch. 2019. Sciurus niger niger (southern fox squirrel) density and the
dirunal patterns, ocupancy, and detection of sympatric southern fox squirrels and S. carolinensis (eastern gray squirrle) on Spring
Island, South Carolina. Southeastern Naturalist 18:321-333.
Wineland, S. M., R. F. Arrick, S. M. Welch, T. K. Pauley, J. Mosher, J.J. Apodaca, M. Olszack, J. N. Holmes, and J. L. Waldron. 2019.
Environmental DNA (eDNA) improves eastern hellbender (Cryptobranchus alleganiensis alleganiensis) detection over conventional
sampling methods. Environmental DNA 1:86-96.
Wineland, S. M.*, S. M. Welch, T. K. Pauley, J. Mosher, J. J. Apodaca, M. Olszack, J. Holmes, and J. L. Waldron. 2019. Using
environmental DNA (eDNA) and occupancy modelling to identify drivers of eastern hellbender (Cryptobranchus allegaiensis alleganiensis)
extirpation. Freshwater Biology 64:208-221.
Heres, B*, S. M. Welch, and J. L. Waldron. 2018. The effect of mast availability on Eastern Diamondback Rattlesnake (Crotalus
adamanteus) ambush site selection. Southeastern Naturalist 17:117-129.
Andrews, K. A., and J. L. Waldron. 2017. Comparative overwintering ecology of a coastal and inland population of canebrake
rattlesnakes (Crotalus horridus) in the southern United States. Pp. 196-212 in Dreslik, M. J., W. K. Hayes, S. J. Beaupre, and S. P.
Mackessy (eds.), The Biology of Rattlesnakes II. ECO Herpetological Publishing and Distribution, Rodeo, New Mexico.
Edwards, E.*, T. K. Pauley, and J. L. Waldron. 2016. Estimating Gyrinophilus porphyriticus detection probability using
multiple methods. Journal of Herpetology 50:126-129.
Fill, J. M.*, W. J. Platt, S. M. Welch, J. L. Waldron, T. A. Mousseau. 2015. Updating models for restoration and management of fiery
ecosystems. Forest Ecology and Management 356:54-63.
Fill, J. M.*, J. L. Waldron, S. M. Welch, M. Martin*, J. Cantrell, Stephen H. Bennett, W. Kalinowsky, J. Holloway, and T. Mousseau. 2015.
Breeding and reproductive phenology of eastern diamond-backed rattlesnake (Crotalus adamanteus) in South Carolina. Journal of
Herpetology 49:570-573.
Fill, J. M.*, J. L. Waldron, S. M. Welch, J. W. Gibbons, S. H. Bennett, and T. A. Mousseau. 2015. Using multiscale spatial models to assess
potential surrogate habitat for an imperiled reptile. PLoS ONE. 10(4): e0123307. DOI: 10.1371/journal.pone.0123307
Oswald, H. R.*, J. L. Waldron, S. M. Welch, and T. A. Mousseau. 2015. Environmental effects on southern two-lined salamander (Eurycea
cirrigera) nest-site selection. Copeia 103:7-13.
Fill, J. M.*, S. M. Welch, H. Brown, J. L. Waldron, A. S. Weakley, and T. A. Mousseau. 2014. Life-history correlates of plant endemism in
longleaf pine ecosystems. Southeastern Naturalist 13:484-492.
Waldron, J. L., S. M. Welch, S. H. Bennett, W. G. Kalinowsky, and T. A. Mousseau. 2013. Life History Constraints Contribute to the
Vulnerability of a Declining North American Rattlesnake. Biological Conservation 159:530-538.
Waldron, J. L, S. M. Welch, J. Holloway, and T. A. Mousseau. 2013. Using Occupancy Models to Examine Human-Wildlife Interactions.
Human Dimensions of Wildlife 18:138-151.
Fill, J. M.*, S. M. Welch, J. L. Waldron, and T. A. Mousseau. 2012. The reproductive response of an endemic bunchgrass indicates historical
timing of a keystone process. Ecosphere 3(7), Article 61; http://www.esajournals.org/doi/pdf/10.1890/ES12-00044.1.
Bennett, S. H., J. L. Waldron, and S. M. Welch. 2012. Light bait improves capture success of aquatic funnel trap sampling for larval
amphibians. Southeastern Naturalist: 11:49-58.
Waldron, J. L., S. M. Welch, and S. H. Bennett. 2008. Vegetation structure and the habitat specificity of a declining North American reptile:
a remnant of former landscapes. Biological Conservation 141:2477-2482.
Welch, S. M., J. L. Waldron, A. G. Eversole, and J. C. Simose. 2008. Seasonal Variation and Ecological Effects of Camp Shelby Burrowing
Crayfish (Fallicambarus gordoni) Burrows. American Midland Naturalist 159:378-384.
Waldron, J. L. and T. K. Pauley. 2007. Green Salamander (Aneides aeneus) growth and age at reproductive maturity. Journal of
Herpetology 41:638-644.
Waldron, J. L., et al. 2006. Habitat specificity and home-range size as attributes of species vulnerability to extinction: A case study using
sympatric rattlesnakes. Animal Conservation 9:414-420.
Waldron, J. L., J. D. Lanham, and S. H. Bennett. 2006. Using behaviorally-based seasons to investigate canebrake rattlesnake (Crotalus
horridus) movement patterns and habitat selection. Herpetologica 62:391-400.
Waldron, J. L., and W. J. Humphries. 2005. Arboreal habitat use by the green salamander in South Carolina. Journal of Herpetology
39:486-492.
Waldron, J. L., C. K. Dodd Jr., and J. D. Corser. 2003. Leaf litterbags: Factors affecting capture of stream-dwelling salamanders. Applied
Herpetology 1:123-36.
Waldron, J. L., Z. Felix, W. J. Humphries, and T. K. Pauley. 2001. Herpetofauna of the Bluestone National Wild and Scenic River, West
Virginia. West Virginia Academy of Science 73:1-10.
Journal of Wildlife Management.
Kelley, A.*, S. M. Welch, J. Holloway, J.W. Dillman, A. Atkinson, and J. L. Waldron. 2022. Effectiveness of long-distance translocation of
Eastern Diamondback Rattlesnakes. Wildlife Society Bulletin e1291. Https://doi.org/10.1002/wsb.1291.
Jungen, M.*, Z. Ross*, J. Cooley*, M. Martin, J. Holloway, S. M. Welch, and J. L. Waldron. 2019. Monitoring eastern diamondback
rattlesnakes using a novel external radio-transmitter attachment method. Copeia 107:411-416.
Amspacher, K.*, B. Bauer, J. Waldron, E. Wiggers, and S. Welch. 2019. Sciurus niger niger (southern fox squirrel) density and the
dirunal patterns, ocupancy, and detection of sympatric southern fox squirrels and S. carolinensis (eastern gray squirrle) on Spring
Island, South Carolina. Southeastern Naturalist 18:321-333.
Wineland, S. M., R. F. Arrick, S. M. Welch, T. K. Pauley, J. Mosher, J.J. Apodaca, M. Olszack, J. N. Holmes, and J. L. Waldron. 2019.
Environmental DNA (eDNA) improves eastern hellbender (Cryptobranchus alleganiensis alleganiensis) detection over conventional
sampling methods. Environmental DNA 1:86-96.
Wineland, S. M.*, S. M. Welch, T. K. Pauley, J. Mosher, J. J. Apodaca, M. Olszack, J. Holmes, and J. L. Waldron. 2019. Using
environmental DNA (eDNA) and occupancy modelling to identify drivers of eastern hellbender (Cryptobranchus allegaiensis alleganiensis)
extirpation. Freshwater Biology 64:208-221.
Heres, B*, S. M. Welch, and J. L. Waldron. 2018. The effect of mast availability on Eastern Diamondback Rattlesnake (Crotalus
adamanteus) ambush site selection. Southeastern Naturalist 17:117-129.
Andrews, K. A., and J. L. Waldron. 2017. Comparative overwintering ecology of a coastal and inland population of canebrake
rattlesnakes (Crotalus horridus) in the southern United States. Pp. 196-212 in Dreslik, M. J., W. K. Hayes, S. J. Beaupre, and S. P.
Mackessy (eds.), The Biology of Rattlesnakes II. ECO Herpetological Publishing and Distribution, Rodeo, New Mexico.
Edwards, E.*, T. K. Pauley, and J. L. Waldron. 2016. Estimating Gyrinophilus porphyriticus detection probability using
multiple methods. Journal of Herpetology 50:126-129.
Fill, J. M.*, W. J. Platt, S. M. Welch, J. L. Waldron, T. A. Mousseau. 2015. Updating models for restoration and management of fiery
ecosystems. Forest Ecology and Management 356:54-63.
Fill, J. M.*, J. L. Waldron, S. M. Welch, M. Martin*, J. Cantrell, Stephen H. Bennett, W. Kalinowsky, J. Holloway, and T. Mousseau. 2015.
Breeding and reproductive phenology of eastern diamond-backed rattlesnake (Crotalus adamanteus) in South Carolina. Journal of
Herpetology 49:570-573.
Fill, J. M.*, J. L. Waldron, S. M. Welch, J. W. Gibbons, S. H. Bennett, and T. A. Mousseau. 2015. Using multiscale spatial models to assess
potential surrogate habitat for an imperiled reptile. PLoS ONE. 10(4): e0123307. DOI: 10.1371/journal.pone.0123307
Oswald, H. R.*, J. L. Waldron, S. M. Welch, and T. A. Mousseau. 2015. Environmental effects on southern two-lined salamander (Eurycea
cirrigera) nest-site selection. Copeia 103:7-13.
Fill, J. M.*, S. M. Welch, H. Brown, J. L. Waldron, A. S. Weakley, and T. A. Mousseau. 2014. Life-history correlates of plant endemism in
longleaf pine ecosystems. Southeastern Naturalist 13:484-492.
Waldron, J. L., S. M. Welch, S. H. Bennett, W. G. Kalinowsky, and T. A. Mousseau. 2013. Life History Constraints Contribute to the
Vulnerability of a Declining North American Rattlesnake. Biological Conservation 159:530-538.
Waldron, J. L, S. M. Welch, J. Holloway, and T. A. Mousseau. 2013. Using Occupancy Models to Examine Human-Wildlife Interactions.
Human Dimensions of Wildlife 18:138-151.
Fill, J. M.*, S. M. Welch, J. L. Waldron, and T. A. Mousseau. 2012. The reproductive response of an endemic bunchgrass indicates historical
timing of a keystone process. Ecosphere 3(7), Article 61; http://www.esajournals.org/doi/pdf/10.1890/ES12-00044.1.
Bennett, S. H., J. L. Waldron, and S. M. Welch. 2012. Light bait improves capture success of aquatic funnel trap sampling for larval
amphibians. Southeastern Naturalist: 11:49-58.
Waldron, J. L., S. M. Welch, and S. H. Bennett. 2008. Vegetation structure and the habitat specificity of a declining North American reptile:
a remnant of former landscapes. Biological Conservation 141:2477-2482.
Welch, S. M., J. L. Waldron, A. G. Eversole, and J. C. Simose. 2008. Seasonal Variation and Ecological Effects of Camp Shelby Burrowing
Crayfish (Fallicambarus gordoni) Burrows. American Midland Naturalist 159:378-384.
Waldron, J. L. and T. K. Pauley. 2007. Green Salamander (Aneides aeneus) growth and age at reproductive maturity. Journal of
Herpetology 41:638-644.
Waldron, J. L., et al. 2006. Habitat specificity and home-range size as attributes of species vulnerability to extinction: A case study using
sympatric rattlesnakes. Animal Conservation 9:414-420.
Waldron, J. L., J. D. Lanham, and S. H. Bennett. 2006. Using behaviorally-based seasons to investigate canebrake rattlesnake (Crotalus
horridus) movement patterns and habitat selection. Herpetologica 62:391-400.
Waldron, J. L., and W. J. Humphries. 2005. Arboreal habitat use by the green salamander in South Carolina. Journal of Herpetology
39:486-492.
Waldron, J. L., C. K. Dodd Jr., and J. D. Corser. 2003. Leaf litterbags: Factors affecting capture of stream-dwelling salamanders. Applied
Herpetology 1:123-36.
Waldron, J. L., Z. Felix, W. J. Humphries, and T. K. Pauley. 2001. Herpetofauna of the Bluestone National Wild and Scenic River, West
Virginia. West Virginia Academy of Science 73:1-10.