Publications from the Marshall University Herpetology & Applied Conservation Lab
.* denotes student author
Edwards, E.*, B. Mosher, T.K. Pauley, S.M. Welch, and J.L. Waldron. In Review. The effect of liming on spring salamander abundance.
Journal of Wildlife Management.
Gray, E.R.* and Waldron, J.L. In Press. Crotalus adamanteus (Eastern Diamondback Rattlesnake). Male-Male Combat. Herpetological Review.
Gray, E.R.*, Waldron, J.L., Wilkinson, S.L., Miller, M.C., Wilkinson, W., and Holloway, J.H. In Press. Crotalus adamanteus (Eastern Diamondback Rattlesnake). Rain-Harvesting. Herpetological Review.
Kelley, A.*, S. M. Welch, J. Holloway, J.W. Dillman, A. Atkinson, and J. L. Waldron. 2022. Effectiveness of long-distance translocation of
Eastern Diamondback Rattlesnakes. Wildlife Society Bulletin e1291. Https://doi.org/10.1002/wsb.1291.
Martin, M., J. Sealy, T. Ivy, and J.L. Waldron. 2021. Status of the Timber Rattlesnake, Crotalus horridus, in South Carolina. In: The Timber
Rattlesnake: Life History, Distribution, Status, and Conservation Plan. Partners in Amphibian and Reptile Conservation.
Kelley, A.G.*, J.L. Waldron, S.M. Welch, J. Cooley, R. Garniewicz, and J.D. Holloway. 2020. Crotalus adamanteus (Eastern Diamond-backed
Rattlesnake). Long-distance Movement. Herpetological Review 51:345.
DODPARC. 2020. Recommended Best Management Practices for the Eastern Diamond-backed Rattlesnake on Department of Defense
Installations. DoD Legacy Resource Management Program. https://www.denix.osd.mil/dodparc/parc-resources/index.html.
Jungen, M.*, Z. Ross*, J. Cooley*, M. Martin, J. Holloway, S. M. Welch, and J. L Waldron. 2019. Monitoring eastern diamondback
rattlesnakes using a novel external radio-transmitter attachment method. Copeia 107:411-416.
Amspacher, K.*, B. Bauer, J. Waldron, E. Wiggers, and S. Welch. 2019. Sciurus niger niger (southern fox squirrel) density and the
diurnal patterns, occupancy, and detection of sympatric southern fox squirrels and S. carolinensis (eastern gray squirrel) on Spring
Island, South Carolina. Southeastern Naturalist 18:321-333.
Wineland, S. M., R. F. Arrick, S. M. Welch, T. K. Pauley, J. Mosher, J.J. Apodaca, M. Olszack, J. N. Holmes, and J. L. Waldron. 2019.
Environmental DNA (eDNA) improves eastern hellbender (Cryptobranchus alleganiensis alleganiensis) detection over conventional
sampling methods. Environmental DNA 1:86-96.
Wineland, S. M.*, S. M. Welch, T. K. Pauley, J. Mosher, J. J. Apodaca, M. Olszack, J. Holmes, and J. L. Waldron. 2019. Using environmental
DNA (eDNA) and occupancy modelling to identify drivers of Eastern Hellbender (Cryptobranchus alleganiensis alleganiensis) extirpation.
Freshwater Biology 64:208-221.
Bonisoli-Alquati, A., S. Ostermiller, D. E. Beasley, S. M. Welch, A. P. Møller, and T. A. Mousseau. 2018. Faster development covaries with
higher DNA damage in grasshoppers (Chorthippus albomarginatus) from Chernobyl. Phsyiological and Biochemical Zoology 91:776-787.
Meshaka, W. E. Jr., T. K. Pauley, J. Russell, and K. M. Herrick*. 2018. Life history traits in the northern ring-necked snake, Diadophis
punctatus edwardsii (Merrem, 1820), in West Virginia. The Herpetological Bulletin 144: 14018.
Heres, B*, S. M. Welch, and J. L. Waldron. 2018. The effect of mast availability on Eastern Diamondback Rattlesnake (Crotalus
adamanteus) ambush site selection. Southeastern Naturalist. Southeastern Naturalist 17:117-129.
Waldron, J.L. 2017. Review of America's Snake: The Rise and FAll of the Timber Rattlesnake, by Ted Levin, Herpetological Review 48:234-235.
Andrews, K. A., and J. L. Waldron. 2017. Comparative overwintering ecology of a coastal and inland population of canebrake
rattlesnakes (Crotalus horridus) in the southern United States. Pp. 196-212 in Dreslik, M. J., W. K. Hayes, S. J. Beaupre, and S. P.
Mackessy (eds.), The Biology of Rattlesnakes II. ECO Herpetological Publishing and Distribution, Rodeo, New Mexico.
Edwards, E.*, T. K. Pauley, and J. L. Waldron. 2016. Estimating Gyrinophilus porphyrticius detection probability using multiple methods.
Journal of Herpetology 50:126-129.
Fill, J. M.*, W. J. Platt, S. M. Welch, J. L. Waldron, T. A. Mousseau. 2015. Updating models for restoration and management of fiery
ecosystems. Forest Ecology and Management 356:54-63.
Fill, J. M.*, J. L. Waldron, S. M. Welch, M. Martin*, J. Cantrell, Stephen H. Bennett, W. Kalinowsky, J. Holloway, and T. Mousseau. 2015.
Breeding and reproductive phenology of eastern diamond-backed rattlesnakes (Crotalus adamanteus) in South Carolina. Journal of
Herpetology 49:570-573.
Fill, J. M.*, J. L. Waldron, S. M. Welch, J. W. Gibbons, S. H. Bennett, and T. A. Mousseau. 2015. Using multiscale spatial models to assess
potential surrogate habitat for an imperiled reptile. PLoS ONE 10(4):e0123307. DOI: 10.1371/journal.pone.0123307
Oswald, H. R.*, J. L. Waldron, S. M. Welch, and T. A. Mousseau. 2015. Environmental effects on southern two-lined salamander (Eurycea
cirrigera) nest-site selection. Copeia 103:7-13.
Fill, J. M.*, S. M. Welch, H. Brown, J. L. Waldron, A. S. Weakley, and T. A. Mousseau. 2014. Life-history correlates of plant endemism in
longleaf pine ecosystems. Southeastern Naturalist 13:484-492.
Kroschel, W. A.*, W. B. Sutton, C. J. W. McClure, and T. K. Pauley. 2014. Decline of the Cheat Mountain Salamander over a 32-Year Period
and the Potential Influence of Competition from a Sympatric Species. Journal of Herpetology 48:415-422.
Waldron, J. L., S. M. Welch, S. H. Bennett, W. G. Kalinowsky, and T. A. Mousseau. 2013. Life History Constraints Contribute to the
Vulnerability of a Declining North American Rattlesnake. Biological Conservation 159:530-538.
Waldron, J. L, S. M. Welch, J. Holloway, and T. A. Mousseau. 2013. Using Occupancy Models to Examine Human-Wildlife Interactions.
Human Dimensions of Wildlife 18:138-151.
Fill, J. M.*, S. M. Welch, J. L. Waldron, and T. A. Mousseau. 2012. The reproductive response of an endemic bunchgrass indicates historical
timing of a keystone process. Ecosphere 3(7), Article 61; http://www.esajournals.org/doi/pdf/10.1890/ES12-00044.1.
Bennett, S. H., J. L. Waldron, and S. M. Welch. 2012. Light bait improves capture success of aquatic funnel trap sampling for larval
amphibians. Southeastern Naturalist: 11:49-58.
Palis, J.G., J.L. Waldron, S.H. Bennett, W.G. Kalinowsky. J.N. Holmes, Z.S. Barfield, M.N. Mimier, and A.P. Dahl. 2012. Crotalus adamanteus
(Eastern Diamondback Rattlesnake). Reproduction/February and March breeding. Herpetological Review 43:657-658.
Waldron, J. L., S. M. Welch, and S. H. Bennett. 2008. Vegetation structure and the habitat specificity of a declining North American reptile:
a remnant of former landscapes. Biological Conservation 141:2477-2482.
Welch, S. M., J. L. Waldron, A. G. Eversole, and J. C. Simose. 2008. Seasonal Variation and Ecological Effects of Camp Shelby Burrowing
Crayfish (Fallicambarus gordoni) Burrows. American Midland Naturalist 159:378-384.
Pauley, T. K. 2008. The Appalachian inferno: historical causes for the disjunct distribution of Plethodon nettingi (Cheat Mountain
Salamander). Northeastern Naturalist 15:595–606.
Pauley, T. K. 2008. Status of the federally threatened Cheat Mountain Salamander, Plethodon nettingi (Amphibia: Caudata): sixty years
later. Proceedings of the West Virginia Academy of Science 80:26–29.
Barry, D.*, T. K. Pauley, and J. Maerz. 2008. Amphibian use of man-made ponds on forest clear-cuts in the Allegheny Mountains. Applied
Herpetology 5:121-128.
Pauley, T. K. 2008. The Appalachian Inferno: A possible reason for the disjunct distribution of a federally protected salamander species,
Plethodon nettingi (Cheat Mountain Salamander). Northeastern Naturalist 15:595-606.
Pauley, B. A. and T. K. Pauley. 2007 Survey of Abandoned Coals Mines for Amphibians and Reptiles in New River Gorge National River,
West Virginia. Proceedings of the West Virginia Academy of Science 79:22-30.
Pauley, T. K. 2007. Revised notes on the range of the Cheat Mountain Salamander, Plethodon nettingi, (Amphibia: Caudata). Proceedings
of the West Virginia Academy of Science 79:16-21.
Waldron, J. L. and T. K. Pauley. 2007. Green Salamander (Aneides aeneus) growth and age at reproductive maturity. Journal of
Herpetology 41:638-644.
Rainwater, T.R., J.L. Waldron, S.M. Welch, S.G. Platt. 2007. Agkistrodon piscivorus piscivorus (Eastern Cottonmouth). Predation.
Herpetological Review 38:202.
Wooten, J.* and T. K. Pauley. 2007. Sexual Size Dimorphism in Five Species of Dusky Salamanders (Caudata: Plethodontidae:
Desmognathus) from West Virginia. Proceedings of the West Virginia Academy of Science 79:9-15.
Wooten, J.* and T. K. Pauley. 2007. Amphibians of the upland habitats within the New River Gorge National River, West Virginia.
Proceedings of the West Virginia Academy of Science 79:32-44.
Felix, Z.* and T. K. Pauley. 2006. Diets of sympatric Black Mountain and Seal Salamanders. Northeastern Naturalist 13:469-476.
Sutton, W. R.*, K. E. Rastall, and T. K. Pauley. 2006. Diet Analysis and feeding strategies of Rana pipiens in a West Virginia wetland.
Herpetological Review 37:152-153.
Waldron, J. L., et al. 2006. Habitat specificity and home-range size as attributes of species vulnerability to extinction: A case study using
sympatric rattlesnakes. Animal Conservation 9:414-420.
Waldron, J. L., J. D. Lanham, and S. H. Bennett. 2006. Using behaviorally-based seasons to investigate canebrake rattlesnake (Crotalus
horridus) movement patterns and habitat selection. Herpetologica 62:391-400.
Watson M.B. and T.K. Pauley. 2006. Spatial Distribution of Turtles in the Great Kanawha River, West Virginia. Proceedings of the West
Virginia Academy of Science 78:14-25.
Humphries, W. J. and T. K. Pauley. 2005. Life History of the Hellbender, Cryptobranchus alleganiensis, in a West Virginia Stream.
American Midland Naturalist 154:135-142.
Waldron, J. L., and W. J. Humphries. 2005. Arboreal habitat use by the green salamander in South Carolina. Journal of Herpetology
39:486-492.
Raimondo, S.*, T. K. Pauley, and L. Butler. 2003. Potential impacts of Bacillus thruingiensis var. kurstaki on five salamander species in
West Virginia. Northeastern Naturalist 10: 25-38.
Waldron, J. L., C. K. Dodd Jr., and J. D. Corser. 2003. Leaf litterbags: Factors affecting capture of stream-dwelling salamanders. Applied
Herpetology 1:123-36.
Brophy, T. R.* and T. K. Pauley. 2002. Reproduction in West Virginia Populations of the Southern Two-lined Salamander (Eurycea cirrigera).
Maryland Naturalist 45:13–22.
Felix, Z.* and T. K. Pauley. 2002. Distribution and Habitat of Desmognathus welteri, Black Mountain Salamander, in West Virginia.
Proceedings of the West Virginia Academy of Science 74:1-5.
Smith, K.G., J.L. Waldron, and W.J. Barichivich. 2002. Elaphe obsoleta obsoleta (black rat snake). Predation. Herpetological Review
33:213-214.
Pauley, T. K., J. C. Mitchell, and M. B. Watson. 2001. The first reported population of the erythristic morph of Plethodon cinereus in
Pennsylvania. Northeastern Naturalist 8:355-358.
Waldron, J. L.*, Z. Felix, W. J. Humphries, and T. K. Pauley. 2001. Herpetofauna of the Bluestone National Wild and Scenic River, West
Virginia. West Virginia Academy of Science 73:1-10.
Barichivich, W.J., K.G. Smith, and J.L. Waldron. 2001. Life history notes: Gyrinophilus porphyriticus danielsi (Blue Ridge Spring Salamander).
Behavior. Herpetological Review 32:177-178.
Humphries, W. H.* and T. K. Pauley. 2000. Seasonal changes in nocturnal activity of the hellbender, Cryptobranchus a. alleganiensis in West
Virginia. Journal of Herpetology 34:604-607.
Pauley, T. K. 2000. Amphibians and reptiles in wetland habitats of West Virginia. Proceedings of the West Virginia Academy of Science
72:78-88.
Humphries, W. J.*, Z. I. Felix*, W. H. Martin, T. K. Pauley.1999. A new lizard species, Cnemidophorus s. sexlineatus, in West Virginia.
Proceedings of the West Virginia Academy of Science 71:1-3.
Mitchell, J. C., T. K. Pauley, D. L. Withers, S. M. Roble, B. T. Miller, A. L. Braswell, P. V. Cupp, and C. S. Hobson.1999. Conservation Status of
The Southern Appalachian Herpetofauna. Virginia Journal of Science 50:1-35.
Pauley, T. K. and M. Little. 1998. New Technique to Monitor Larval and Juvenile Salamanders in Stream Habitats. Banisteria 12:32-36.
Brophy, T. R.* and T. K. Pauley. 1997. Dietary Comparison of Eurycea cirrigera (Southern Two-lined Salamander) Larvae from Pond and
Stream Habitats in Southern West Virginia. Proceedings of the West Virginia Academy of Science 69:4-10.
Jewell, R. D. and T. K. Pauley. 1995. Notes on the Reproductive Biology of the salamander Plethodon richmondi (Nettingi and Mittleman) in
West Virginia. Herpetological Natural History, 3:91-93.
Pauley, T. K. and E. C. Keller, Jr. 1993. Relationships of Burrow Size, Soil Temperature, and Body Size and Temperature in the Salamanders
Plethodon wehrlei and Plethodon cinereus. Proceedings of the West Virginia Academy of Science 65:2-7.
Canterbury, R. C. and T. K. Pauley. 1994. Time of Mating and Egg Deposition of West Virginia Populations of the Green Salamander,
Aneides aeneus (Cope and Packard). Journal of Herpetology, 28:431-434.
Joy, J. E., T. K. Pauley, and M. L. Little. 1993. Prevalence and intensity of Thelandros magnavulvaris and Omeia papillocauda (Nematoda) in
two species of Desmognathine salamanders from West Virginia. Journal of Helminthol. Soc. Wash. 60:93-95.
Canterbury, R. C. and T. K. Pauley. 1990. Food Habits of Aneides aeneus in West Virginia. Proceedings of the West Virginia Academy of
Science 62:47-50.
Pauley, T. K. 1986. Herpetology Field Notes: Two Record Size Salamanders. Proceeding of the West Virginia Academy of Science
58:59-60.
Pauley, T. K. 1986. Additional notes on the Range and Distribution of the Cheat Mountain Salamander, Plethodon nettingi. Proceedings of
the West Virginia Academy Of Science 58:56-59.
Little, M. L. and T. K. Pauley. 1986. A New Record of the Diploid Species of Gray Treefrog, Hyla chrysoscelis. Proceedings of the West
Virginia Academy of Science 58:57-59.
Pauley, T. K. 1986. Amphibians and reptiles of the 1985 foray. The Redstart 53:34-35.
Pauley, T. K, 1983. West Virginia Dusky Salamanders. The Restart 50:85-91.
Pauley, T. K. 1984. The amphibians and reptiles of the 1984 Brooks Bird Club Foray. he Restart 52:40-42.
Pauley, T. K. 1982. Saving the salamander. The Highlands Voice 14:5.
Pauley, T. K. 1981. The range and distribution of the Cheat Mountain salamander, Plethodon nettingi. Proceedings of the West Virginia
Academy of Science 53:31-35.
Pauley, T. K. 1980. Field notes on the distribution of terrestrial amphibians and reptiles of the West Virginia mountains above 970 meters.
Proceedings of the West Virginia Academy of Science 52:84-92.
Pauley, T. K. 1979. The status of the woodland salamanders in West Virginia. The Restart 46:106-112.
Pauley, T. K. 1978. Temperature as a single factor regulating Plethodon habitat partitioning. Proceedings of the West Virginia Academy of Science 50:77-83.
Pauley, T. K. 1978. Moisture as a factor regulating habitat partitioning between two sympatric Plethodon species. Journal of Herpetology
12:491-493.
Pauley, T. K. 1978. Food types and distribution as a Plethodon habitat partitioning factor. Bulletin of the Maryland Herpetological Society
14:79-82.
Pauley, T. K. 1978. Plants as indicators of occurrence of two sympatric Plethodon species. Bulletin of the Maryland Herpetological Society
14:29-35.
Pauley, T. K. 1977. An Unusual Date for a Sandhill Crane in West Virginia. The Restart 41:109.
Pauley, T. K. 1974. A leucistic Plethodon cinereus from West Virginia. The Restart
Pauley, T. K. 1973. The Status of the Genus Carphophis in Ohio and West Virginia. Proceedings of the West Virginia Academy of Science
45:64-70.
Pauley, T. K. and W. H. England. 1969. Time of Mating and Egg Deposition in the salamander, Plethodon wehrlei, Fowler and Dunn, in West
Virginia. Proceedings of the West Virginia Academy of Science 45:64-70.
Edwards, E.*, B. Mosher, T.K. Pauley, S.M. Welch, and J.L. Waldron. In Review. The effect of liming on spring salamander abundance.
Journal of Wildlife Management.
Gray, E.R.* and Waldron, J.L. In Press. Crotalus adamanteus (Eastern Diamondback Rattlesnake). Male-Male Combat. Herpetological Review.
Gray, E.R.*, Waldron, J.L., Wilkinson, S.L., Miller, M.C., Wilkinson, W., and Holloway, J.H. In Press. Crotalus adamanteus (Eastern Diamondback Rattlesnake). Rain-Harvesting. Herpetological Review.
Kelley, A.*, S. M. Welch, J. Holloway, J.W. Dillman, A. Atkinson, and J. L. Waldron. 2022. Effectiveness of long-distance translocation of
Eastern Diamondback Rattlesnakes. Wildlife Society Bulletin e1291. Https://doi.org/10.1002/wsb.1291.
Martin, M., J. Sealy, T. Ivy, and J.L. Waldron. 2021. Status of the Timber Rattlesnake, Crotalus horridus, in South Carolina. In: The Timber
Rattlesnake: Life History, Distribution, Status, and Conservation Plan. Partners in Amphibian and Reptile Conservation.
Kelley, A.G.*, J.L. Waldron, S.M. Welch, J. Cooley, R. Garniewicz, and J.D. Holloway. 2020. Crotalus adamanteus (Eastern Diamond-backed
Rattlesnake). Long-distance Movement. Herpetological Review 51:345.
DODPARC. 2020. Recommended Best Management Practices for the Eastern Diamond-backed Rattlesnake on Department of Defense
Installations. DoD Legacy Resource Management Program. https://www.denix.osd.mil/dodparc/parc-resources/index.html.
Jungen, M.*, Z. Ross*, J. Cooley*, M. Martin, J. Holloway, S. M. Welch, and J. L Waldron. 2019. Monitoring eastern diamondback
rattlesnakes using a novel external radio-transmitter attachment method. Copeia 107:411-416.
Amspacher, K.*, B. Bauer, J. Waldron, E. Wiggers, and S. Welch. 2019. Sciurus niger niger (southern fox squirrel) density and the
diurnal patterns, occupancy, and detection of sympatric southern fox squirrels and S. carolinensis (eastern gray squirrel) on Spring
Island, South Carolina. Southeastern Naturalist 18:321-333.
Wineland, S. M., R. F. Arrick, S. M. Welch, T. K. Pauley, J. Mosher, J.J. Apodaca, M. Olszack, J. N. Holmes, and J. L. Waldron. 2019.
Environmental DNA (eDNA) improves eastern hellbender (Cryptobranchus alleganiensis alleganiensis) detection over conventional
sampling methods. Environmental DNA 1:86-96.
Wineland, S. M.*, S. M. Welch, T. K. Pauley, J. Mosher, J. J. Apodaca, M. Olszack, J. Holmes, and J. L. Waldron. 2019. Using environmental
DNA (eDNA) and occupancy modelling to identify drivers of Eastern Hellbender (Cryptobranchus alleganiensis alleganiensis) extirpation.
Freshwater Biology 64:208-221.
Bonisoli-Alquati, A., S. Ostermiller, D. E. Beasley, S. M. Welch, A. P. Møller, and T. A. Mousseau. 2018. Faster development covaries with
higher DNA damage in grasshoppers (Chorthippus albomarginatus) from Chernobyl. Phsyiological and Biochemical Zoology 91:776-787.
Meshaka, W. E. Jr., T. K. Pauley, J. Russell, and K. M. Herrick*. 2018. Life history traits in the northern ring-necked snake, Diadophis
punctatus edwardsii (Merrem, 1820), in West Virginia. The Herpetological Bulletin 144: 14018.
Heres, B*, S. M. Welch, and J. L. Waldron. 2018. The effect of mast availability on Eastern Diamondback Rattlesnake (Crotalus
adamanteus) ambush site selection. Southeastern Naturalist. Southeastern Naturalist 17:117-129.
Waldron, J.L. 2017. Review of America's Snake: The Rise and FAll of the Timber Rattlesnake, by Ted Levin, Herpetological Review 48:234-235.
Andrews, K. A., and J. L. Waldron. 2017. Comparative overwintering ecology of a coastal and inland population of canebrake
rattlesnakes (Crotalus horridus) in the southern United States. Pp. 196-212 in Dreslik, M. J., W. K. Hayes, S. J. Beaupre, and S. P.
Mackessy (eds.), The Biology of Rattlesnakes II. ECO Herpetological Publishing and Distribution, Rodeo, New Mexico.
Edwards, E.*, T. K. Pauley, and J. L. Waldron. 2016. Estimating Gyrinophilus porphyrticius detection probability using multiple methods.
Journal of Herpetology 50:126-129.
Fill, J. M.*, W. J. Platt, S. M. Welch, J. L. Waldron, T. A. Mousseau. 2015. Updating models for restoration and management of fiery
ecosystems. Forest Ecology and Management 356:54-63.
Fill, J. M.*, J. L. Waldron, S. M. Welch, M. Martin*, J. Cantrell, Stephen H. Bennett, W. Kalinowsky, J. Holloway, and T. Mousseau. 2015.
Breeding and reproductive phenology of eastern diamond-backed rattlesnakes (Crotalus adamanteus) in South Carolina. Journal of
Herpetology 49:570-573.
Fill, J. M.*, J. L. Waldron, S. M. Welch, J. W. Gibbons, S. H. Bennett, and T. A. Mousseau. 2015. Using multiscale spatial models to assess
potential surrogate habitat for an imperiled reptile. PLoS ONE 10(4):e0123307. DOI: 10.1371/journal.pone.0123307
Oswald, H. R.*, J. L. Waldron, S. M. Welch, and T. A. Mousseau. 2015. Environmental effects on southern two-lined salamander (Eurycea
cirrigera) nest-site selection. Copeia 103:7-13.
Fill, J. M.*, S. M. Welch, H. Brown, J. L. Waldron, A. S. Weakley, and T. A. Mousseau. 2014. Life-history correlates of plant endemism in
longleaf pine ecosystems. Southeastern Naturalist 13:484-492.
Kroschel, W. A.*, W. B. Sutton, C. J. W. McClure, and T. K. Pauley. 2014. Decline of the Cheat Mountain Salamander over a 32-Year Period
and the Potential Influence of Competition from a Sympatric Species. Journal of Herpetology 48:415-422.
Waldron, J. L., S. M. Welch, S. H. Bennett, W. G. Kalinowsky, and T. A. Mousseau. 2013. Life History Constraints Contribute to the
Vulnerability of a Declining North American Rattlesnake. Biological Conservation 159:530-538.
Waldron, J. L, S. M. Welch, J. Holloway, and T. A. Mousseau. 2013. Using Occupancy Models to Examine Human-Wildlife Interactions.
Human Dimensions of Wildlife 18:138-151.
Fill, J. M.*, S. M. Welch, J. L. Waldron, and T. A. Mousseau. 2012. The reproductive response of an endemic bunchgrass indicates historical
timing of a keystone process. Ecosphere 3(7), Article 61; http://www.esajournals.org/doi/pdf/10.1890/ES12-00044.1.
Bennett, S. H., J. L. Waldron, and S. M. Welch. 2012. Light bait improves capture success of aquatic funnel trap sampling for larval
amphibians. Southeastern Naturalist: 11:49-58.
Palis, J.G., J.L. Waldron, S.H. Bennett, W.G. Kalinowsky. J.N. Holmes, Z.S. Barfield, M.N. Mimier, and A.P. Dahl. 2012. Crotalus adamanteus
(Eastern Diamondback Rattlesnake). Reproduction/February and March breeding. Herpetological Review 43:657-658.
Waldron, J. L., S. M. Welch, and S. H. Bennett. 2008. Vegetation structure and the habitat specificity of a declining North American reptile:
a remnant of former landscapes. Biological Conservation 141:2477-2482.
Welch, S. M., J. L. Waldron, A. G. Eversole, and J. C. Simose. 2008. Seasonal Variation and Ecological Effects of Camp Shelby Burrowing
Crayfish (Fallicambarus gordoni) Burrows. American Midland Naturalist 159:378-384.
Pauley, T. K. 2008. The Appalachian inferno: historical causes for the disjunct distribution of Plethodon nettingi (Cheat Mountain
Salamander). Northeastern Naturalist 15:595–606.
Pauley, T. K. 2008. Status of the federally threatened Cheat Mountain Salamander, Plethodon nettingi (Amphibia: Caudata): sixty years
later. Proceedings of the West Virginia Academy of Science 80:26–29.
Barry, D.*, T. K. Pauley, and J. Maerz. 2008. Amphibian use of man-made ponds on forest clear-cuts in the Allegheny Mountains. Applied
Herpetology 5:121-128.
Pauley, T. K. 2008. The Appalachian Inferno: A possible reason for the disjunct distribution of a federally protected salamander species,
Plethodon nettingi (Cheat Mountain Salamander). Northeastern Naturalist 15:595-606.
Pauley, B. A. and T. K. Pauley. 2007 Survey of Abandoned Coals Mines for Amphibians and Reptiles in New River Gorge National River,
West Virginia. Proceedings of the West Virginia Academy of Science 79:22-30.
Pauley, T. K. 2007. Revised notes on the range of the Cheat Mountain Salamander, Plethodon nettingi, (Amphibia: Caudata). Proceedings
of the West Virginia Academy of Science 79:16-21.
Waldron, J. L. and T. K. Pauley. 2007. Green Salamander (Aneides aeneus) growth and age at reproductive maturity. Journal of
Herpetology 41:638-644.
Rainwater, T.R., J.L. Waldron, S.M. Welch, S.G. Platt. 2007. Agkistrodon piscivorus piscivorus (Eastern Cottonmouth). Predation.
Herpetological Review 38:202.
Wooten, J.* and T. K. Pauley. 2007. Sexual Size Dimorphism in Five Species of Dusky Salamanders (Caudata: Plethodontidae:
Desmognathus) from West Virginia. Proceedings of the West Virginia Academy of Science 79:9-15.
Wooten, J.* and T. K. Pauley. 2007. Amphibians of the upland habitats within the New River Gorge National River, West Virginia.
Proceedings of the West Virginia Academy of Science 79:32-44.
Felix, Z.* and T. K. Pauley. 2006. Diets of sympatric Black Mountain and Seal Salamanders. Northeastern Naturalist 13:469-476.
Sutton, W. R.*, K. E. Rastall, and T. K. Pauley. 2006. Diet Analysis and feeding strategies of Rana pipiens in a West Virginia wetland.
Herpetological Review 37:152-153.
Waldron, J. L., et al. 2006. Habitat specificity and home-range size as attributes of species vulnerability to extinction: A case study using
sympatric rattlesnakes. Animal Conservation 9:414-420.
Waldron, J. L., J. D. Lanham, and S. H. Bennett. 2006. Using behaviorally-based seasons to investigate canebrake rattlesnake (Crotalus
horridus) movement patterns and habitat selection. Herpetologica 62:391-400.
Watson M.B. and T.K. Pauley. 2006. Spatial Distribution of Turtles in the Great Kanawha River, West Virginia. Proceedings of the West
Virginia Academy of Science 78:14-25.
Humphries, W. J. and T. K. Pauley. 2005. Life History of the Hellbender, Cryptobranchus alleganiensis, in a West Virginia Stream.
American Midland Naturalist 154:135-142.
Waldron, J. L., and W. J. Humphries. 2005. Arboreal habitat use by the green salamander in South Carolina. Journal of Herpetology
39:486-492.
Raimondo, S.*, T. K. Pauley, and L. Butler. 2003. Potential impacts of Bacillus thruingiensis var. kurstaki on five salamander species in
West Virginia. Northeastern Naturalist 10: 25-38.
Waldron, J. L., C. K. Dodd Jr., and J. D. Corser. 2003. Leaf litterbags: Factors affecting capture of stream-dwelling salamanders. Applied
Herpetology 1:123-36.
Brophy, T. R.* and T. K. Pauley. 2002. Reproduction in West Virginia Populations of the Southern Two-lined Salamander (Eurycea cirrigera).
Maryland Naturalist 45:13–22.
Felix, Z.* and T. K. Pauley. 2002. Distribution and Habitat of Desmognathus welteri, Black Mountain Salamander, in West Virginia.
Proceedings of the West Virginia Academy of Science 74:1-5.
Smith, K.G., J.L. Waldron, and W.J. Barichivich. 2002. Elaphe obsoleta obsoleta (black rat snake). Predation. Herpetological Review
33:213-214.
Pauley, T. K., J. C. Mitchell, and M. B. Watson. 2001. The first reported population of the erythristic morph of Plethodon cinereus in
Pennsylvania. Northeastern Naturalist 8:355-358.
Waldron, J. L.*, Z. Felix, W. J. Humphries, and T. K. Pauley. 2001. Herpetofauna of the Bluestone National Wild and Scenic River, West
Virginia. West Virginia Academy of Science 73:1-10.
Barichivich, W.J., K.G. Smith, and J.L. Waldron. 2001. Life history notes: Gyrinophilus porphyriticus danielsi (Blue Ridge Spring Salamander).
Behavior. Herpetological Review 32:177-178.
Humphries, W. H.* and T. K. Pauley. 2000. Seasonal changes in nocturnal activity of the hellbender, Cryptobranchus a. alleganiensis in West
Virginia. Journal of Herpetology 34:604-607.
Pauley, T. K. 2000. Amphibians and reptiles in wetland habitats of West Virginia. Proceedings of the West Virginia Academy of Science
72:78-88.
Humphries, W. J.*, Z. I. Felix*, W. H. Martin, T. K. Pauley.1999. A new lizard species, Cnemidophorus s. sexlineatus, in West Virginia.
Proceedings of the West Virginia Academy of Science 71:1-3.
Mitchell, J. C., T. K. Pauley, D. L. Withers, S. M. Roble, B. T. Miller, A. L. Braswell, P. V. Cupp, and C. S. Hobson.1999. Conservation Status of
The Southern Appalachian Herpetofauna. Virginia Journal of Science 50:1-35.
Pauley, T. K. and M. Little. 1998. New Technique to Monitor Larval and Juvenile Salamanders in Stream Habitats. Banisteria 12:32-36.
Brophy, T. R.* and T. K. Pauley. 1997. Dietary Comparison of Eurycea cirrigera (Southern Two-lined Salamander) Larvae from Pond and
Stream Habitats in Southern West Virginia. Proceedings of the West Virginia Academy of Science 69:4-10.
Jewell, R. D. and T. K. Pauley. 1995. Notes on the Reproductive Biology of the salamander Plethodon richmondi (Nettingi and Mittleman) in
West Virginia. Herpetological Natural History, 3:91-93.
Pauley, T. K. and E. C. Keller, Jr. 1993. Relationships of Burrow Size, Soil Temperature, and Body Size and Temperature in the Salamanders
Plethodon wehrlei and Plethodon cinereus. Proceedings of the West Virginia Academy of Science 65:2-7.
Canterbury, R. C. and T. K. Pauley. 1994. Time of Mating and Egg Deposition of West Virginia Populations of the Green Salamander,
Aneides aeneus (Cope and Packard). Journal of Herpetology, 28:431-434.
Joy, J. E., T. K. Pauley, and M. L. Little. 1993. Prevalence and intensity of Thelandros magnavulvaris and Omeia papillocauda (Nematoda) in
two species of Desmognathine salamanders from West Virginia. Journal of Helminthol. Soc. Wash. 60:93-95.
Canterbury, R. C. and T. K. Pauley. 1990. Food Habits of Aneides aeneus in West Virginia. Proceedings of the West Virginia Academy of
Science 62:47-50.
Pauley, T. K. 1986. Herpetology Field Notes: Two Record Size Salamanders. Proceeding of the West Virginia Academy of Science
58:59-60.
Pauley, T. K. 1986. Additional notes on the Range and Distribution of the Cheat Mountain Salamander, Plethodon nettingi. Proceedings of
the West Virginia Academy Of Science 58:56-59.
Little, M. L. and T. K. Pauley. 1986. A New Record of the Diploid Species of Gray Treefrog, Hyla chrysoscelis. Proceedings of the West
Virginia Academy of Science 58:57-59.
Pauley, T. K. 1986. Amphibians and reptiles of the 1985 foray. The Redstart 53:34-35.
Pauley, T. K, 1983. West Virginia Dusky Salamanders. The Restart 50:85-91.
Pauley, T. K. 1984. The amphibians and reptiles of the 1984 Brooks Bird Club Foray. he Restart 52:40-42.
Pauley, T. K. 1982. Saving the salamander. The Highlands Voice 14:5.
Pauley, T. K. 1981. The range and distribution of the Cheat Mountain salamander, Plethodon nettingi. Proceedings of the West Virginia
Academy of Science 53:31-35.
Pauley, T. K. 1980. Field notes on the distribution of terrestrial amphibians and reptiles of the West Virginia mountains above 970 meters.
Proceedings of the West Virginia Academy of Science 52:84-92.
Pauley, T. K. 1979. The status of the woodland salamanders in West Virginia. The Restart 46:106-112.
Pauley, T. K. 1978. Temperature as a single factor regulating Plethodon habitat partitioning. Proceedings of the West Virginia Academy of Science 50:77-83.
Pauley, T. K. 1978. Moisture as a factor regulating habitat partitioning between two sympatric Plethodon species. Journal of Herpetology
12:491-493.
Pauley, T. K. 1978. Food types and distribution as a Plethodon habitat partitioning factor. Bulletin of the Maryland Herpetological Society
14:79-82.
Pauley, T. K. 1978. Plants as indicators of occurrence of two sympatric Plethodon species. Bulletin of the Maryland Herpetological Society
14:29-35.
Pauley, T. K. 1977. An Unusual Date for a Sandhill Crane in West Virginia. The Restart 41:109.
Pauley, T. K. 1974. A leucistic Plethodon cinereus from West Virginia. The Restart
Pauley, T. K. 1973. The Status of the Genus Carphophis in Ohio and West Virginia. Proceedings of the West Virginia Academy of Science
45:64-70.
Pauley, T. K. and W. H. England. 1969. Time of Mating and Egg Deposition in the salamander, Plethodon wehrlei, Fowler and Dunn, in West
Virginia. Proceedings of the West Virginia Academy of Science 45:64-70.
